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3-fluorotyrosine carbonising tissue colorbearer conchoclination Cornelia Otis Skinner dictyosome (perroncito 1910) doyle-scribner rule George Gills Ra. global feature measure hot load strength integral square-error method Keimax Matukituki R. methyl propenyl ketone parazoan pre-orogenic reducing resister sea surface observation speed of test zero discharge power