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Anthocephalus bawshere biofunctionalization carsounds direct-attached Dubiecko glyptophidium lucidum gnathocephalon high-ash content Ludlow Bone Bed Mammaria Nehruvian palpebral fissure (or lid-slit) parallel deal Pearce Syzygium rehderianum T-shaped rigid frame bridge TcHHW time selector within-family selection